The Nationals, who trailed 9-1 after two innings and 10-2 after four, completed their biggest comeback win in nearly 12 years.
Uggla, who made the club as a non-roster invitee in the spring, went 3-for-4 with five RBIs against the Braves, the club that released him last June.
The homer, his first of the season, came against Braves closer Jason Grilli (0-1), who was trying to earn his eighth save.
After falling behind 4-0 after two innings, the Jays knocked Clay Buchholz out with a five-run third and never trailed.
Jose Bautista drove in three runs, one with his fifth homer of the season. Josh Donaldson had three hits, including a solo homer, and two RBIs.
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